Rune Trees

Yggdrasil in the village: Glory buys runes, and every rune counts in every run after.

Glory. Every run pays 1 a depth gained, 2 a node cleared and 25 a boss; every class level pays 10 more. Glory is per class and per season.

The tree. Each class has its own: a free heart, and five branches round it. A branch runs trunk → fork → two arms; between them a wheel of small runes, a notable opposite the way in, and in the middle the branch's keystone. Bridges join each branch to the next, so the arms make a web.

Buying a rune needs one beside it you own. You can refund a rune if everything else still links to the heart, or all of them at once. Runes are bought between runs only.

Berserker · Elementalist · Huntress · Grovekeeper

Berserker

Berserker's Heart · 75 runes · 2080 Glory for all of them

Runes

Rune Lore ×2
Trunk · 8 each
Skills recharge 5% faster, each
Rune-Carver
Fork · 30
Skills recharge 6% faster · +6% skill duration
Wide Wyrd
Arm · 12
+4% area
Wide Wyrd
Arm end · 40
+12% area
Long Saga
Arm · 12
+5% skill duration
Long Saga
Arm end · 40
+12% skill duration
Galdr ×5
Wheel · 15 each
Skills recharge 3% faster / +3% area / +4% skill duration / Skills recharge 3% faster / +3% area
Runic Echo
Notable · 50
Skills recharge 6% faster · +6% skill duration
The Fourth Rune
Keystone · 150
Every fourth skill you cast is cast again. Whirlwind, Skin-Changer and Valkyrie's Refusal are never cast twice (nor counted).
Battle-Runes
Bridge · 15
Skills recharge 3% faster · +3% attack speed

The Axe

Honed Edge ×2
Trunk · 8 each
+8% weapon damage, each
Wide Cleave
Fork · 30
+15% melee damage · +5% area
Quick Hands
Arm · 12
+5% attack speed
Quick Hands
Arm end · 40
+12% attack speed
Brutal Blows
Arm · 12
+10% critical damage
Brutal Blows
Arm end · 40
+25% critical damage
Smith's Craft ×5
Wheel · 15 each
+5% weapon damage / +4% attack speed / +8% critical damage / +4% attack speed / +5% weapon damage
Axe-Storm
Notable · 50
+8% attack speed · +8% weapon damage
ARSENAL
Keystone · 120
Carry more than one weapon: [E] on a weapon puts it in a free skill slot, where it attacks on its own (and adds its bonuses).
Blood on the Blade
Bridge · 15
+3% weapon damage · +5% Rage gained

Hide & Charge

Thick Hide ×2
Trunk · 8 each
+8 life, each
Restless Charge
Fork · 30
Charge recharges 20% faster
Rage-Born
Arm · 12
+8% Rage gained
Rage-Born
Arm end · 40
+25% Rage gained
Fleet Foot
Arm · 12
+4% move speed
Fleet Foot
Arm end · 40
+8% move speed · Charge recharges 15% faster
Bear-Skin ×5
Wheel · 15 each
+6 life / 2% less damage taken / +3% move speed / 2% less damage taken / +6 life
Bear's Hide
Notable · 50
8% less damage taken · +10 life
Bloodthirst
Keystone · 120
3% of damage dealt heals you
Wanderer
Bridge · 15
+3% move speed · +3% warden's light radius

Luck & Light

Fortune ×2
Trunk · 8 each
+5% luck (more drops), each
Lantern Soul
Fork · 30
+12% warden's light radius
Keen Eye
Arm · 12
+2% critical chance
Keen Eye
Arm end · 40
+6% critical chance
Magpie
Arm · 12
+10% pickup reach
Magpie's Eye
Arm end · 40
+25% pickup reach · +6% luck (more drops)
Norn-Thread ×5
Wheel · 15 each
+4% warden's light radius / +4% experience / +2% critical chance / +4% experience / +4% warden's light radius
Mímir's Well
Notable · 50
+12% experience
Loki's Luck
Keystone · 120
+25% luck (more drops) · +4% critical chance
Omens
Bridge · 15
+4% luck (more drops) · +2% damage

The Gods' Favours

Devotion ×2
Trunk · 8 each
+3% damage, each
Tyr's Favour
Fork · 30
+15% physical damage · +100% physical skill drops
Bloodletting
Arm · 12
+6% bleed damage
Hel's Favour
Arm end · 40
+15% bleed damage · +100% bleed skill drops
Keen Eye
Arm · 12
+6% projectile damage
Ullr's Favour
Arm end · 40
+15% projectile damage · +100% projectile skill drops
Offering ×5
Wheel · 15 each
+3% damage / +5% Rage gained / +6% summon damage / +5% Rage gained / +6% summon damage
Odin's Favour
Notable · 50
+20% Rage gained · +10% area damage · +60% area skill drops
Fenrir's Kin
Keystone · 120
+25% summon damage · +150% summon skill drops
Skald's Verse
Bridge · 15
+4% skill duration · +2% damage

Elementalist

Elementalist's Heart · 75 runes · 2080 Glory for all of them

Muspel

Kindling ×2
Trunk · 8 each
+5% fire damage, each
Muspel's Favour
Fork · 30
+15% fire damage · +100% fire skill drops
Wildfire
Arm · 12
+4% area
Wildfire
Arm end · 40
+12% area
Long Burn
Arm · 12
+5% skill duration
Long Burn
Arm end · 40
+12% skill duration
Cinder ×5
Wheel · 15 each
+4% fire damage / +3% area / +8% critical damage / +3% area / +4% fire damage
Surtr's Brand
Notable · 50
+10% fire damage · +10% critical damage
Sons of Muspel
Keystone · 120
+20% fire damage · +15% warden's light radius
Steam
Bridge · 15
+3% fire damage · +3% frost damage

Niflheim

Hoarfrost ×2
Trunk · 8 each
+5% frost damage, each
Niflheim's Favour
Fork · 30
+15% frost damage · +100% frost skill drops
Rime-Shell
Arm · 12
+5% Ward
Rime-Shell
Arm end · 40
+12% Ward
Deep Cold
Arm · 12
+5% skill duration
Deep Cold
Arm end · 40
+12% skill duration
Ice-Vein ×5
Wheel · 15 each
+4% frost damage / +5 life / 2% less damage taken / +5 life / +4% frost damage
Glacier's Heart
Notable · 50
6% less damage taken · +8 life
Fimbulwinter
Keystone · 120
+20% frost damage · +20% Ward
Hailstorm
Bridge · 15
+3% frost damage · +3% lightning damage

Thor

Spark ×2
Trunk · 8 each
+5% lightning damage, each
Thor's Favour
Fork · 30
+15% lightning damage · +100% lightning skill drops
Quick Cast
Arm · 12
+5% attack speed
Quick Cast
Arm end · 40
+12% attack speed
Thunderclap
Arm · 12
+2% critical chance
Thunderclap
Arm end · 40
+6% critical chance
Storm-Charge ×5
Wheel · 15 each
+4% lightning damage / +3% attack speed / +2% critical chance / +3% attack speed / +4% lightning damage
Thunderhead
Notable · 50
+10% lightning damage · +12% critical damage
Forked Lightning
Keystone · 120
+1 chains · +10% lightning damage
Static
Bridge · 15
+3% lightning damage · +5% Resonance gained

Resonance

Hum ×2
Trunk · 8 each
+6% Resonance gained, each
Deep Resonance
Fork · 30
+12% Resonance gained · +10% Discharge power
Hoarder
Arm · 12
+5% Resonance gained
Hoarder
Arm end · 40
+1 to the Resonance cap · +4% damage
Breaker
Arm · 12
+6% Discharge power
Breaker
Arm end · 40
+1 to the Resonance cap · +15% Discharge power
Echo ×5
Wheel · 15 each
+4% Discharge power / +4% Resonance gained / Skills recharge 3% faster / +4% Resonance gained / +4% Discharge power
Standing Wave
Notable · 50
+1 to the Resonance cap · +10% Discharge power
Overtone
Keystone · 150
+2 to the Resonance cap · +20% Discharge power
Attunement
Bridge · 15
+5% Resonance gained · Skills recharge 3% faster

Seiðr-craft

Wand-Lore ×2
Trunk · 8 each
+8% weapon damage, each
Staff-Craft
Fork · 30
+12% weapon damage · +5% attack speed
Blink-Walker
Arm · 12
Blink recharges 8% faster
Blink-Walker
Arm end · 40
Blink recharges 20% faster · +4% move speed
Völva's Ward
Arm · 12
+5% Ward
Völva's Ward
Arm end · 40
+12% Ward · +8 life
Seiðr-Thread ×5
Wheel · 15 each
+5% weapon damage / Skills recharge 3% faster / +5 life / Skills recharge 3% faster / +5% weapon damage
Spell-Weaver
Notable · 50
Skills recharge 6% faster · +8% weapon damage
MANY WANDS
Keystone · 120
Carry more than one wand or staff: [E] on a weapon puts it in a free skill slot, where it casts on its own (and adds its bonuses).
Burning Stave
Bridge · 15
+3% weapon damage · +3% fire damage

Huntress

Huntress's Heart · 75 runes · 2080 Glory for all of them

The Bow

Fletching ×2
Trunk · 8 each
+8% weapon damage, each
True Flight
Fork · 30
+12% projectile damage · +5% attack speed
Quick Draw
Arm · 12
+5% attack speed
Quick Draw
Arm end · 40
+12% attack speed
Bodkin Heads
Arm · 12
+6% projectile damage
Bodkin Heads
Arm end · 40
Arrows pierce +1 enemy · +8% projectile damage
Bowyer's Craft ×5
Wheel · 15 each
+5% weapon damage / +3% attack speed / +4% projectile damage / +3% attack speed / +5% weapon damage
Twin Nock
Notable · 50
+1 projectiles
MANY BOWS
Keystone · 120
Carry more than one bow: [E] on a weapon puts it in a free skill slot, where it shoots on its own (and adds its bonuses).
Barbed Heads
Bridge · 15
+3% projectile damage · +3% bleed damage

The Hunt

Keen Sight ×2
Trunk · 8 each
+2% critical chance, each
Marked Prey
Fork · 30
+4% critical chance · +8% skill duration
Deadeye
Arm · 12
+10% critical damage
Deadeye
Arm end · 40
+25% critical damage
Open Wound
Arm · 12
+6% bleed damage
Open Wound
Arm end · 40
+15% bleed damage · +100% bleed skill drops
Tracker ×5
Wheel · 15 each
+2% critical chance / +4% bleed damage / +8% critical damage / +4% bleed damage / +2% critical chance
Blood Trail
Notable · 50
+10% bleed damage · +10% skill duration
Skadi's Quarry
Keystone · 120
+6% critical chance · +25% critical damage
Baited Snare
Bridge · 15
+3% bleed damage · +3% trap damage

The Wilds

Woodcraft ×2
Trunk · 8 each
+5% luck (more drops), each
Trapper
Fork · 30
+15% trap damage · +100% trap skill drops
Iron Jaws
Arm · 12
+6% trap damage
Iron Jaws
Arm end · 40
+12% trap damage · +8% area
Falconer
Arm · 12
+6% beast damage
Falconer
Arm end · 40
+15% beast damage · +100% beast skill drops
Forager ×5
Wheel · 15 each
+4% luck (more drops) / +3% area / +8% pickup reach / +3% area / +4% luck (more drops)
Wide Country
Notable · 50
+12% area · +15% pickup reach
Wild Kin
Keystone · 120
+25% beast damage · +150% beast skill drops · +10% luck (more drops)
Game Trail
Bridge · 15
+4% luck (more drops) · +3% move speed

Fleet Foot

Light Step ×2
Trunk · 8 each
+3% move speed, each
Quick Roll
Fork · 30
Tumble recharges 20% faster
Hard to Pin
Arm · 12
+5 life
Hard to Pin
Arm end · 40
+10 life · 4% less damage taken
Wind-Runner
Arm · 12
+4% move speed
Wind-Runner
Arm end · 40
+8% move speed · Tumble recharges 15% faster
Hare's Leap ×5
Wheel · 15 each
+3% move speed / Tumble recharges 6% faster / +5 life / Tumble recharges 6% faster / +3% move speed
Hare's Heart
Notable · 50
Tumble recharges 15% faster · +5% move speed
THIRD WIND
Keystone · 150
+1 Tumble charge
Running Start
Bridge · 15
+3% move speed · +5% Momentum gained

Momentum

Stride ×2
Trunk · 8 each
+6% Momentum gained, each
Keen Wind
Fork · 30
+12% Momentum gained · +10% Hawk's Eye duration
Unbroken Stride
Arm · 12
+5% Momentum gained
Unbroken Stride
Arm end · 40
+10% Momentum gained · +4% attack speed
Long Glide
Arm · 12
+6% Hawk's Eye duration
Long Glide
Arm end · 40
+15% Hawk's Eye duration
Updraft ×5
Wheel · 15 each
+4% Hawk's Eye duration / +4% Momentum gained / +3% attack speed / +4% Momentum gained / +4% Hawk's Eye duration
High Circling
Notable · 50
+12% Hawk's Eye duration · +8% Momentum gained
UNSHAKEN
Keystone · 120
A blow that lands leaves you 50% of your Momentum, instead of none
Loosed in Stride
Bridge · 15
+5% Momentum gained · +3% weapon damage

Grovekeeper

Grovekeeper's Heart · 75 runes · 2080 Glory for all of them

The Pack

Fang and Claw ×2
Trunk · 8 each
+8% beast damage, each
Pack Leader
Fork · 30
+12% beast damage · Beasts bite 5% faster
Quick Jaws
Arm · 12
Beasts bite 5% faster
Quick Jaws
Arm end · 40
Beasts bite 12% faster
Old Friends
Arm · 12
+6% summon damage
Old Friends
Arm end · 40
+12% summon damage · +100% beast skill drops
Den-Mates ×5
Wheel · 15 each
+5% beast damage / Beasts bite 3% faster / +4% summon damage / Beasts bite 3% faster / +5% beast damage
Alpha
Notable · 50
+15% beast damage · Beasts bite 8% faster
A FULL DEN
Keystone · 150
+1 of each beast at your side
Forest Folk
Bridge · 15
+3% summon damage · +3% area

The Grove

Green Thumb ×2
Trunk · 8 each
+6% burst power, each
Ripe Pods
Fork · 30
+12% burst power · +5% area
Spore Cloud
Arm · 12
+5% area
Spore Cloud
Arm end · 40
+12% area · +100% area skill drops
Splinters
Arm · 12
+6% burst power
Splinters
Arm end · 40
+15% burst power
Seedbed ×5
Wheel · 15 each
+4% burst power / +3% area / +4% summon damage / +3% area / +4% burst power
Second Planting
Notable · 50
+1 sprout
WILDWOOD
Keystone · 120
+1 sprout · +25% burst power
Thorn Pods
Bridge · 15
+3% burst power · +3% bleed damage

Thorn & Root

Whittling ×2
Trunk · 8 each
+8% weapon damage, each
Briar Hand
Fork · 30
+12% weapon damage · +8% bleed damage
Bramble
Arm · 12
+6% bleed damage
Bramble
Arm end · 40
+15% bleed damage · +100% bleed skill drops
Deep Roots
Arm · 12
+6% skill duration
Deep Roots
Arm end · 40
+12% skill duration · +6% attack speed
Hedgecraft ×5
Wheel · 15 each
+5% weapon damage / +4% bleed damage / +3% attack speed / +4% bleed damage / +5% weapon damage
Strangleroot
Notable · 50
+10% bleed damage · +10% skill duration
THORNHEART
Keystone · 120
+25% weapon damage · +25% bleed damage
Ironbark Switch
Bridge · 15
+3% weapon damage · +3 life

Bark

Thick Bark ×2
Trunk · 8 each
+6 life, each
Heartwood
Fork · 30
+12 life · 4% less damage taken
Mossback
Arm · 12
+8% Ward
Mossback
Arm end · 40
+20% Ward · 3% less damage taken
Tunneler
Arm · 12
Burrow recharges 6% faster
Tunneler
Arm end · 40
Burrow recharges 15% faster · +4% move speed
Growth Rings ×5
Wheel · 15 each
+5 life / +5% Ward / 2% less damage taken / Burrow recharges 5% faster / +5 life
Old Stump
Notable · 50
+15 life · 5% less damage taken
OLD AS OAK
Keystone · 120
+30 life · 8% less damage taken · +25% Ward
Sheltering Boughs
Bridge · 15
+3 life · +5% Kinship gained

Kinship

Fellowship ×2
Trunk · 8 each
+6% Kinship gained, each
Many Voices
Fork · 30
+12% Kinship gained · +10% Stampede duration
Shared Bread
Arm · 12
+5% Kinship gained
Shared Bread
Arm end · 40
+10% Kinship gained · +5 life
Thundering Hooves
Arm · 12
+6% Stampede duration
Thundering Hooves
Arm end · 40
+15% Stampede duration
Hearth-Song ×5
Wheel · 15 each
+4% Stampede duration / +4% Kinship gained / +3% summon damage / +4% Kinship gained / +4% Stampede duration
The Wild Hunt
Notable · 50
+12% Stampede duration · +8% Kinship gained
NEVER ALONE
Keystone · 120
Your friends can take 14% more of each blow meant for you
Call of Kin
Bridge · 15
+5% Kinship gained · +3% beast damage