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The season
Everyone starts from nothing in a new season: levels, Glory, the Rune Tree, the Delver, hats and materials, and a new map. The season that ends is kept (its boards too); a player in a run finishes it first. It cannot be taken back from here.
Every season
Resetting a board deletes that season's runs (the Live and Deepest boards); class levels come from the saves and stay.